The epic tale of the meek Neptune, Compa, and IF on their search for Histoire.
Along the way, Neptune will bump into three other buxom female characters named Noire, Vert, and Blanc in Gamindustri. These three characters are known as the three goddesses, or CPUs. Upon transformation, they become Black/White/Green Heart which are their true goddess forms. Each has their own particular personality traits and flaws, with the subtle references to the actual game consoles they personify.
As for the villain in the game, she is known as Arfoire, who is hell-bent on wreaking havoc in Gamindustri. She is the embodiment of all things evil and wrong in the gaming world and will constantly appear to thwart you in your journey to find Histoire.
The world itself is divided into the four major areas of Planeptune, Leanbox, Lastation and Lowee. Within each are countless dungeons for exploration and questing. Between the dungeons and quests, you will frequent places called Basilicoms where you meet other quirky characters to help drive the storyline and keep things interesting as you progress through the game.